In Kentucky just after the Civil War, the Hayden-Colby feud leads to Jed Colby being sent to prison for 15 years for murder. The Haydens head for Nevada and when Colby gets out of prison he heads there also seeking revenge. The head of the Hayden family tries to avoid more killing but the inevitable showdown has to occur, complicated by Lynn Hayden and Ellen Colby's plans to marry.
Clint Belmet (Gary Cooper) is a bit of a firebrand and is sentenced to at least 30 days in jail, but his partners, Bill Jackson (Ernest Torrence) and Jim Bridger (Tully Marshall) talk a sympathetic Frenchwoman named Felice (Lili Damita) into telling the bumbling, drunken marshal that Clint had married her the previous night. Clint is released so he can accompany Felice on the wagon train heading west to California.
Western pardners Jeff and Cash find a baby boy in an otherwise deserted emigrants' camp, and clash over which is to be "father." They are still bitterly feuding years later when they own adjacent ranches. Bill, the foundling whom Cash has raised to young manhood, wants to end the feud and extends an olive branch toward Jeff, who now has a lovely daughter. But during a mining venture, the bitterness escalates. Is Bill to be set against his own adoptive father?
With little luck at keeping a job in the city a New Yorker tries work in the country and eventually finds his way leading a herd of cattle to the West Coast.
The earliest surviving film featuring Lon Chaney in a major role, By the Sun's Ray's was but one of several 2-reel westerns starring the florid Murdock MacQuarrie. MacQuarrie plays a detective investigating a series of gold shipment robberies. Along the way, he falls for a mine superintendent's pretty daughter (Agnes Vernon), much to the dismay of a sullen mine office clerk (Chaney), who is also smitten with the girl...